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Name: Dr. Samiullah Khan
Designation: Professor
Department: Microbiology


Qualifications: Ph.D (Sweden)
Postdoctorate (Sweden)


Phone: +92-51 9064-3011
Email: samikhan@qau.edu.pk
Status: On Job
Other Weblink: https://www.qau.edu.pk/profile.php?id=805020

View Resume

Research Interests Publications Conferences Research Projects

          Bacteriophages and their Enzyme as Enzy biotics for the control of multi drug resistance bacteria

·         Microbial production of industrially important metabolites with special focus on enzymes and antimicrobials

·         Special research focuses is on hydrolase group of enzyme e.g. glycosidase from extremophiles, Proteases and Phytases

·         Probiotics and prebiotics

·         Divesity, Cultivation, Screening and Characterization of edible and non-edible mushrooms of Phamaceutical Importance

·         Use of Algae for Biotechnological application e.g. Algae in Food Biotechnology, Pharmaceutical Biotechnology and paper and pulp industry

·         Screening and isolation of glycosidases, Proteases and Phytases producing bacteria and fungi from species adapted to the extreme environments.

·         Cloning of the gene that encodes the enzyme and overexpression of the selected genes.

·          Purification and characterized of the expressed genes (Enzymes)

·         Bioinformatics, Structure/Function studies and  modelling of enzymes

·         Enzymes from thermophiles (activity, kinetics and deactivation/denaturation studies)

·         Cultivation of microorganisms (recombinant E. coli)

·         Enzyme engineering and designing of new biocatalyst

·         Enzymatic production of prebiotic Resistant Starch

·         Enzymes production for Bio-refinery such as conversion of chicken feather wastes into value added products

·         Enzymatic modification of natural products such as antioxidants for use in different industrial products e.g. food, pharmaceuticals and cosmetics using synthetic approach.


1         Nawaz A, Khalid NA, Zaheer S, Khan MI, Khalid A,  Shah AA, Badshah M and Khan S. Isolation and Characterization of a Potent Bacteriophage KA Targeting an Antibiotic-Resistant Human Pathogenic Strain of Klebsiella pneumoniae KP1. Microbail Pathogenesis 2025 Article No. YMPAT_108150. https://doi.org/10.1016/j.micpath.2025.108150.  (I.F 3.5)

 

2         Masood H, Duarte A. S. R., Akbar A, Abbas S. H, Badshah M,. Shah AA and  Khan S. The assessment of antibiotics usage and disease trends in commercial broiler farms of Potohar region in Pakistan: Implications for food safety. Animal and Plant Sciences 2025. 35(6) (2025). 605 (I.F 0.60)

 

3         Nawaz A, Khalid NA, Majid A, Shahzadi M, Zafar S, Khan MI, Khan S, Khalid A,  Shah AA, Badshah M and Khan S. Characterization of bacteriophages PAA and PAM and evaluation of their antibiotic synergy against Pseudomonas aeruginosa PAZMYU isolated from urine sample. BMC Microbiology 2025, Article No. 605. https://doi.org/10.1186/s12866-025-04195-x. (I.F 4.2)

 

4         Khan S, Ilyas N,  Nawaz A, Khan MI,  Aamir M, Khalid N.A, Shah AA, Zada NS, Shabbir G, Badshah M and Khan S. Bacteriophage-based Vaccination Platform for Antigen Discovery and Development of Next-Generation Vaccines. Next Research 2025, Article No. 100776 https://doi.org/10.1016/j.nexres.2025.100776 .

 

5         Khan A.Z, Din Ud S, Sharif M,  Hasan F, Khan S, Badshah M, Belduz A.O and Shah AA .Isolation and Characterization of Aspergillus niger AZ2 from Pulp and Paper Mill Effluent that Degrade Lignin under Alkaline Conditions. Current Microbiology. 2025, 82(6): article No 421. https://doi.org/10.1007/s00284-025-04413-9. (I.F 2.4)

 

6         Alam T, Din Ud S, Abdullah M, Ali M, Badshah M, Farman M, Khan S, Hasan F and Shah A.A. Bioactive Metabolites from Radioresistant Bacterium Kocuria sp. TMM 11 and Their Role in Prevention of Ultraviolet-Induced Photodamages. Current Microbiology. 2025, 82(6): article No 243. DOI10.1007/s00284-025-04194-1. (I.F 2.4)

 

7         Din Ud S, Israr A, Jabeen S, Rana Q, Din Ud S, Kun Z, Khan S, Hasan F, Badshah M, Zhou Y.J, and Shah A.A.  Production of Free Fatty Acids from Alkaline Organosolvent Pretreated Sugarcane Bagasse and Trash by Yeast Pichia pastoris. Journal of Waste and Biomass Valorization, 2025. https://doi.org/10.1007/s12649-025-02963-4. (I.F 2.6)

 

8         Din Ud S, Alam T, Khan S, Hasan F, Badshah M, and Shah A.A.  Bioactive metabolites from radioresistant bacterium Kocuria sp. TMM 11 and their role in prevention of Ultraviolet-induced photodamages. Journal of Current Microbiology, 2025. 82:243 (2025). https://doi.org/10.1007/s00284-025-04194-1  (I.F 2.3)

 

9         Zada NS, Belduz A.O, Aleesa H.A, Guler HI, KaraoÄŸlan M, Badshah M, Shah AA, Kalsoom A and Khan S Comparison of active site mutations at subsite +2 of Anoxybacillus ayderensis A9 β-glucosidase for hydrolysis of pNPG and polydatin. BMC Biotechnology, 2025, 25:52 (2025). https://doi.org/10.1186/s12896-025-00984-4

 (IF: 3.2).

 

10      Saba M, Farooq S, Aleesa H.A, Bektas K.I, Belduz A.O, Khan A.Z, Shah AA, Badshah M and Khan S.  Green synthesis of silver nanoparticles using Keratinase from Pseudomonas aeruginosa-C1M, characterization and applications as novel multifunctional biocatalyst. BMC Biotechnology, 2025, 25:27. https://doi.org/10.1186/s12896-025-00959-5  (IF: 3.2).

11      Mansoor A, Mansoor E, Hussain K and Khan S. Clinical efficacy of novel biogenically fabricated titania nanoparticles enriched mouth wash in treating the tooth dentine hypersensitivity: A randomized clinical trial. Pakistan Journal of Medical Sciences, 2025, 41(6):1743-1748. (IF: 1.7).

12      Kalsoom, Din Ud S, Ceylan E, Hasan F, Khan S, çanakçi S, Belduz A.O and Shah AA. Cloning and expression of chromate reductase from Bacillus paramycoides S48 for chromium remediation. Scientific Report, 2025, 15: 18796 (2025). https://doi.org/10.1038/s41598-025-03412-x   (IF: 3.8).

110. Shoukat M, Rehman A, Khan A, Ali I, Nazish M, Abashr F.T, Alrefaei A.F, Khan S, Shah A.A and Badshah M. Enhancement of Biogas Production through Phase Separated Anaerobic Co-Digestion of Cattle Manure with Fruit and Vegetable Waste. Polish Journal of Environmental Studies, 2025.

DOI: https://doi.org/10.15244/pjoes/201937  (IF: 1.4).

 

14      Khan M.A.N, Mishqat A, Bukhari S.M.A.U.S, Haq A, Ali M, Khan S, Shah A.A, Ali N, Ahmed S and Badshah M. Effect of different Parameters on Biogas Production during Next Generation Digestion, Anaerobic Digestion Coupled with Microbial Electrolysis Cell. Renewable Energy.2025, 243:122620. https://doi.org/10.1016/j.renene.2025.122620 (IF: 9.0)

15       Mansoor A,  Mansoor E,  Mansoor E, Shah A,  Asjid U  ∙ João Filipe Brochado Martins J.F.B, Khan S and  Palma P.J.  Synthesis of Novel Titania Nanoparticles using Corn Silky Hair Fibres and their role in developing a Smart Restorative Material in Dentistry. Computational and Structural Biotechnology Journal. 2025, 29: 29-40. (I. F 4.5).

16      Abbas S.H, Khan S, Shah M, Aslam J, Nawaz H, Ilyas N, Gamaryani A, Afridi S.Q, Khan I, Shah B, Shah K, Rashid A, Khan D and Khan S.  Public Health Threats Posed by Biofilms and Innovative Strategies for their Control. Discovery Journal. 2024, Article number   12(3); e197

17      Nawaz A, Zafar S, Alessa AH, Khalid NA, Muqaddas S, Majid A,  Badshah M, Shah AA,  and Khan S. Characterization of ES10 lytic bacteriophage isolated from hospital waste against multidrug-resistant uropathogenic E. coli. Frontiers Microbiology. DOI 10.3389/fmicb.2024.1320974 (I.F 5.2).

 

18      Khan A,  Mehmood K, Nadhman A, Khan S, Shah AA and Shah Z. Microbial Production of Keratinase from Bacillus velezensis strain MAMA: A Novel Enzyme for Eco-Friendly Degradation of Keratin Waste. Heliyon. 2024, 10(12), e32338.  https://doi.org/10.1016/j.heliyon.2024.e32338. (I.F 4.0)

 

19      Ali Q,  Zainab R,  Badshah M, Sarwar W, Khan S, Mustafa G, Ibrahim T and Ahmed S. Prospecting the biodegradation of ciprofloxacin by Stutzerimonas stutzeri R2 and Exiguobacterium indicum strain R4 isolated from pharmaceutical wastewater. H2 Open Journal. (2024), 7(2), 149-162. (I.F 2.1)

 

20      Sharif M, Nawaz A, Naeem S, Saeeda UH, Farid A, Saleh Zada N, Shah AA, Badshah M, Alharbi MT, Tarabulsi MK, Selim S and Khan S. Pharmaceutical and Nutraceutical Importance of Bioactive Metabolites from Macrofungi. Journal of Biological Regulators and Homeostatic Agents. (2024) 38(5), Article No.8019 (I.F 3.2).

 

21      Nawaz A,  Khalid NA, Zafar S, Majid A, Muqaddas S, Saleem S, Shah AA, Badshah M and Khan S. Phage therapy as a revolutionary treatment for multidrug-resistant Pseudomonas aeruginosa infections: A narrative review. The Microbes. 2 (2024): 100030. https://doi.org/10.1016/j.microb.2023.100030

                                                      

22      Fatima A, Ibrahim M, Naseer A, Pervez A, Asad M, Shah AA, Hasan F, Alonazi W.B, Ferheen I and Khan S. Identification, Genome Sequencing, and Characterizations of Helicobacter pylori Sourced from Pakistan. Microorganisms. 2023, 11, 2658. https://doi.org/10.3390/microorganisms11112658  (I.F 4.926)

 

23      Nawaz A,  Zafar S, Muqaddas S, Bukhari S.M.A.S, Khan NA, Shah AA, Badshah M and Khan S. Bacteriophages: an overview of the control strategies against phytopathogens. Egyptian Journal of Biological Pest Control. 2023, 33 Article number: 108 (2023). (I.F 2.3)

 

 

24      Siddiqui S, Khan A, A,  Rahman U, Shah AA, Badshah M, Rehman MM, Hayat A and Khan S. Understanding the Inhibitory Mechanism of Exogenous Protein on Enzymatic Preparation of Resistant Starch. Starch, 2023, Article number 2200264. DOI: 10.1002/star.202200264. (I.F 2.741)

 

25      Haq A, Malik A, Khan A, Weaver E. J, Wang L, Khan H, Khan S, Shah AA, Ahmed S, Jamal A, De Los Reyes F. L and Badshah M. Effect of removal of inhibitors on microbial communities and biogas yield of Jatropha curcas seeds during continuous anaerobic digestion. Journal of Cleaner Production. October 2023, 426(5):139154 (I.F 11.1)

 

26      Khan S.I, Sahinkaya M, Colak D.N, Zada N.S, Uzuner U,  Belduz A.O, Canakci S, Khan A.Z, Khan S, Badshah M and Shah AA. Production and characterization of novel thermostable CotA-laccase from Bacillus altitudinis SL7 and its application for lignin degradation. Enzyme and Microbial Technology, 2024, 172:(110329). (I.F 3.493).

 

27      Khan A, Siddiqui S, Rahman U, Belduz A.O, Shah AA, Badshah M, Hasan F and Khan S.  Enzymatic modification of maize flour improves its functional properties, digestion resistibility, and antioxidant potential. Journal of Food Measurement and Characterization, 2023, 17:6109-6124. (I.F 3.4)

 

28      Saba M, Akhter A, Ahmed H, Mehmood Z, Khan A, Salehzada N, Badshah M, Hasan F, Shah AA, and Khan S.  Sustainable valorization of chicken feathers and grocery waste as organic fertilizer and its impact on yield and quality of Spinach (Spinacia oleracea) plant. Journal of  Communications in Soil Science and Plant Analysis, 2023, 54(21):2995-3005. (I.F 1.8)

 

 

29      Haq A, Rehman M.L, Rana Q.A, Khan A, Sajjad W, Khan H,  Khan S, Shah AA, Hasan F, Ahmed S, Islam A, Badshah M, Shah T.A, Dawoud T.M, and Bourhia M. Production, optimization, and physicochemical characterization of biodiesel from seed oil of indigenously grown Jatropha curcas.

Frontiers in Energy Research, 2023, 11: doi.org/10.3389/fenrg.2023.1225988. (I.F 3.4)

 

30      Rahman A, Ali A, Ahmad F, Ahmad S, Alharbi M, Alasmar A.F, Fayyaz A, Rana Q.A,  Khan S, Hasan F, Badshah M and Shah AA. Unraveling the Radioprotective Mechanisms of UV-Resistant Bacillus subtilis ASM-1 Extracted Compounds through Molecular Docking. Pharmaceuticals, 2023,16(8):1139. (I.F 4.6)

 

31      Sajjad W, Nadeem M, Alam T, Rehman A, Abbasi S.W, Ahmad S, Din S. U,  Khan S, Badshah M, Gul S, Farman M and Shah AA. Biological Evaluation and Computational Studies of Methoxy‒flavones from Newly Isolated Radioresistant Micromonospora aurantiaca Strain TMC‒15. Journal of Applied Biochemistry and Biotechnology, 2023, 195:4915-4935. (I.F 2.431)

 

32      Rahman U, Qasim S, Zada N.S, Ahmad S, Shah AA, Badshah M, Hasan F, and Khan S. Evaluation of Agricultural Wastes as a Sustainable Carbon Source for the Production of β-glucosidase from Bacillus stercoris, its Purification and Characterization. Pakistan Journal of Agricultural Sciences, 2023, 60(2): 367-375. (IF: 1.0).

 

33      Din S. U, Kalsoom, Satti S.M, Uddin S, Mankar S.V, Ceylan E, Hasan F, Khan S, Badshah M, Belduz A.O, çanakçi S, Zhang B, Pasten J.A.L and Shah AA. The Purification and Characterization of a Cutinase-like Enzyme with Activity on Polyethylene Terephthalate (PET) from a Newly Isolated Bacterium Stenotrophomonas maltophilia PRS8 at a Mesophilic Temperature. Applied Sciences, 2023 13(6): 3686. (IF: 2.838)

 

34      Khan A, Ali H, Rahman U, Belduz A.O, Bibi A, Abdurahman  M.A, Shah AA, Badshah M, Hasan F, Kilic  A.O, Ullah A, Jahan S, Rehman MM, Mansoor R and Khan S. Prebiotic potential of enzymatically prepared resistant starch in reshaping gut microbiota and their respond to body physiology. PLOS ONE, 16 May, 2022 (17(5): e0267318). (I.F 3.24)

 

35      Hayat R, Din G, Farooqi A, Khan S, Hasan F, Badshah M and Shah AA. Characterization of melanin pigment from Aspergillus terreus LCM8 and its role in cadmium remediation. International Journal of Environmental Science and Technology, 2023, 20: 3151–3160. (I.F 2.860)

 

36      Arif  M, Kalssom, Shah A.A, Badshah M, Hasan F, Rehman A  and Khan S. Positivity, diagnosis and treatment follow-up of cutaneous leishmaniasis in war-affected areas of Bajaur, Pakistan. Parasitology Research, 2022, 121: 191-198. (I.F 2.403)

 

37      Saba M, Khan A, Ali H, Bibi A, Gul Z, Khan A, Rehman MM, Badshah M, Hasan F, Shah AA and Khan S. Microbial Pretreatment of Chicken Feather and Its Co-digestion With Rice Husk and Green Grocery Waste for Enhanced Biogas Production. Frontiers in Microbiology, 7 April, 2022, 13:792426. (I.F 5.6)

 

38      Mumtaz T, Ahmed N, Hassan N, Badshah M, Khan S and Rehman. Voriconazole nanoparticles-based film forming spray: An efficient approach for potential treatment of topical fungal infections. Journal of Drug Delivery Science and Technology, 2022, 70: (102973). (I.F 3.981)

 

39      Khan S.I, Zarin A, Ahmed S, Hasan F, Belduz A.O, çanakçi S, Khan S, Badshah M, Farman M and  Shah AA. Degradation of lignin by Bacillus altitudinis SL7 isolated from pulp and paper mill effluent. Water Science & Technology, 2022, 85(1): 420 (I.F 1.915)

 

40      Rana Q, Khan M.A.N, Shiekh Z, Parveen S, Ahmed S, Irfan M,  Gauttam R,  Shah AA,  Jamal A, Khan S, and Badshah M. Production of bioethanol and biogas from Spirodela polyrhiza in a biorefinery concept and output energy analysis of the process. Biomass Conversion and Biorefinery, 26 November, 2021. https://doi.org/10.1007/s13399-021-02066-9. (I.F 4.987)

 

41      Zada N.S, Belduz A.O, Güler H.I, Sahinkaya M, Khan S.I, Saba M, Bektas K.I, Kara Y, Kolaylı S, Badshah M, Shah A.A and Khan S. Cloning, Biochemical characterization, and molecular docking of Novel thermostable β-glucosidase BglA9 from Anoxybacillus Ayderensis A9 and its application in de-glycosylation of Polydatin. International Journal of Biological Macromolecules, 2021, 193: 1898-1909. (I.F 6.953)

 

42      Khan A, Akbar S, Okonkwo V, Smith C, , Khan S, Shah AA, Adnan F, Ijaz U.Z,  Ahmed  A and Badshah M. Enrichment of the hydrogenotrophic methanogens for, in-situ biogas up-gradation by recirculation of gases and supply of hydrogen in methanogenic reactor. Bioresource Technology, 2022, 345(2):126219. (I. F. (9.642)

 

43      Zada N.S, Belduz A.O, Guler K.I, Khan A, Sahinkaya M, Kaciran A, Ay H, Badshah M, Shah AA and Khan S. Cloning, expression, biochemical characterization, and Molecular Docking studies of a novel glucose tolerant β-glucosidase from Saccharomonospora sp. NB11. Enzyme and Microbial Technology, August, 2021, 148: (109799). (I.F 3.493).

 

44      Khan S.I, Zada N.S, Sahinkaya M, Colak D.N, Ahmed S, Hasan F, Belduz A.O, Khalil I, Canakci S, Khan S, Badshah M and Shah AA Cloning, expression and biochemical characterization of lignin-degrading DyP-type peroxidase from Bacillus sp. Strain BL5. Enzyme and Microbial Technology, September 2021, 151: (109917). (I.F 3.493).

 

45      Farooqi A, Din G, Hayat R, Badshah M, Khan S and  Shah A.A.Characterization of Bacillus nealsonii strain KBH10 capable of reducing aqueous mercury in laboratory-scale reactor. Water Science & Technology,2021, 83 (9): 2287–2295. (I.F: 1.638).

 

 

46      Haq A,  Mushtaq S, Khan A, Islam A, Khan H, Malik Z.A, Khan S, Shah AA and Badshah M. Evaluation of phytochemical, bioactive, and antifungal potential of Jatropha curcas seed oil and de-oiled seed cake extracts against phytopathogenic fungi. Journal of Plant Pathology, 2021, 103: 863–873. (I.F 1.729).

 

 

47      Khan A, Irfan1 A, Rahman U, Azhar F, Shah1 AA, Badshah M, Hasan F, Rehman F, Malik Z.A, and Khan S. A Ca+2 independent pullulanase from Bacillus licheniformis and its application in the synthesis of resistant starch. Pakistan Journal of Agricultural Sciences, 2021 58(2): 699-709. (IF: 1.0).

 

48      Haq A, Khan A, Malik Z.A, Ahmed M, Khan S, Shah AA, Hasan F and Badshah M. Antimicrobial activities of different parts of two geographically distinct varieties of Jatropha curcas Linn. fruit in Pakistan. Bangladesh Journal of Botany, 2021, 50(2). (IF: 0.35).

 

49      Kalsoom, Batool A, Din G, Din S, Jamil J, Hasan F, Khan S, Badshah M and Shah AA.  Isolation and screening of chromium resistant bacteria from industrial waste for bioremediation purposes. Brazilian Journal of Biology, 2021. (3;83:e242536). https://doi.org/10.1590/1519-6984.242536. (I.F: 1.36).

 

50      Zada N.S, Bari F, Malik Z.A, Bangash F.H, Mansoor R, Shah AA, Badshah M, Khan S. Phenotypic detection of Metallo-β-Lactamase (MBL) in Imipenem-Resistant Pseudomonas aeruginosa, a study from a tertiary care hospital in Peshawar, Pakistan. International Journal of Biosciences, 2021, 18(6):120-128. (IF: 0.553).

 

51      Aijaz M, Malik Z.A, Khan A, Khan S,  Shah AA, Badshah M, Hasan F and Khan S. Antioxidant, Antimicrobial and Cytotoxic Potential of Selected Medicinal Plants Collected from Khanewal Valley, Pakistan. International Journal of Biosciences, 2021, 18(5):86-99. (IF: 0.553).

 

52      Aslam A, Malik Z.A, Akhtar N, Chandra J, Rafique K, Hasan F, Ikram A, Khan S, Badshah M, Ghannoum M.A, and Shah AA. Characterization of biofilms produced by Candida species isolated from tertiary care hospitals, Rawalpindi, Pakistan. International Journal of Biosciences, 2021, 18(4):111-122. (IF: 0.553).

 

53      Arif H.M, Malik Z.A, Irfan M, Hanif F, Hasan F, Khan S, Badshah M and Shah AA. A model for finding new L-asparaginase producing microorganisms using Taguchi design of experiments. International Journal of Biosciences, 2021,18(3), 276-289. (IF: 0.553).

 

54      Ullah A, Malik Z.A, Irfan M, Din S, Rana Q.A, Badshah M, Khan S, Hasan F and Shah AA. Production and characterization of xylanase from Bacillus licheniformis S3 isolated from hot spring. International Journal of Biosciences, 2021,18(4), 144-158. (IF: 0.553).

 

55      Khattak A, Malik Z.A, Haq A, Rana Q.A, Khan H, Rehman F, Khan S, Shah AA and Badshah M. Comparative assessment of acid and enzyme pretreatment of Spirodella polyrhiza for bioethanol production. International Journal of Biosciences, 2021,18(6): 103-119. (IF: 0.553).

 

56      Ali P, Chen F, Hasan H, Sosa A, Khan S, Badshah M and  Shah A.A. Bacterial community characterization of Batura Glacier in the Karakoram Range of Pakistan. International Micrbiology, 2021 (24)183–196. https://doi.org/10.1007/s10123-020-00153-x (I.F 2.479).

 

57      Khan A, Bibi A, Ali H, Rehman A, Qindeel M, Irfan M, Shah AA, Badshah M, Hasan F and Khan S. Development of Resistant Starch Film-coated Microparticles for an Oral Colon-Specific Drug Delivery. Journal of Starch, 2020. https://doi.org/10.1002/star.201900262 (IF: 2.741)

 

58      Khan A, Sidiqqui S, Rahman U, Saba M, Azhar F, Shah AA, Badshah M, Hasan F and Khan S.  Physicochemical properties of resistant starch from maize flour by enzyme treated and its use in cookies formulation. International Journal of food Properties, 2020 23(1) 549-569. (IF: 2.727)

 

59      Haq A,  Adeel S, Khan A, Rana Q, Khan M.A.N, Rafiq M, Ashfaq M, Khan S, Shah A.A, Hasan F, Ahmed S and Badshah M. Screening of lipase producing bacteria and optimization of lipase mediated biodiesel production from Jatropha curcas seed oil using whole cell approach. Bioenergy Research, 2020 https://doi.org/10.1007/s12155-020-10156-1. (IF: 2.814)

 

60      Haq A, Khan A,  Khan H, Khan S, Shah A.A, Hasan F, Ahmed S, Reyes F.L and Badshah M. Enhancement of biogas yield during anaerobic digestion of Jatropha curcas seed by pretreatment and co-digestion with mango peels. Biomass Conversion and Biorefinary 2022, 12: 1595-1603. https://doi.org/10.1007/s13399-020-01064-7. (IF: 4.987)

 

61      Irfan M, Kiran J, Salahuddin, Ameen ullah, Rana Q, Khan S, Hasan F, Badshah M and Shah A.A. Immobilization of β-1,4-xylanase Isolated from Bacillus licheniformis S3. Journal of Basic Microbiology, 2020, 60(7): 600-612. doi: 10.1002/jobm.202000077. (IF: 2.281)

 

62      Chaudry A.A, Mahnoor N, Rahman A, Khan F, Hasan F, Khan S, Badshah M and Shah AA. Antioxidative and radioprotective properties of glycosylated flavonoid, xanthorhamnin from radio-resistant bacterium Bacillus indicus strain TMC-6. Current Microbiology, 2020, 77, 1245-1254. (IF: 2.188)

 

63      Noor H, Satti S.M, Din S, Farman M, Hasan F, Khan S, Badshah M and Shah AA. Insight on esterase from Pseudomonas aeruginosa strain S3 that depolymerize poly (lactic acid) (PLA) at ambient temperature. Polymer Degradation and Stability. 2020,174:109096. (IF: 5.030)

 

64      Rana Q, Khan M.A.N, Irfan M,  Shah AA,  Hasan F, Khan S, Ahmed S, Adnan F, Li W, Ju M and Badshah M. Starved Spirodela polyrhiza and Saccharomyces cerevisiae: A Potent Combination for Sustainable Bioethanol Production. Biomass Conversion and Biorefinery, 2021, 11:1665-1674. https://doi.org/10.1007/s13399-019-00540-z. (IF: 4.987)

 

65      Ali P, Hasan H, Khan S, Badshah M, Shah A.A. Cold-adapted halotolerant Rhodococcus sp. BGI-11, a potential candidate for biotechnological applications. International Journal of Biosciences 2019, 15(2): 475-489. (IF: 0.553).

 

66      Safia B, Hameed S, Khan S, Khan A, Rehmat Y, Rafiq N, Ahmad A, Zakria M. Taxanomy of Micro-invertebrates inhabiting in fresh water algal mats in ponds of Quetta City and Zhob District, Balochistan, Pakistan. International Journal of Biosciences 2019, 15(4): 62-69. (IF: 0.553).

 

67      Din G, Hassan A, Rafiq M, Hasan F, Badshah M, Khan S, Chen G, Ripp S and Shah AA. Characterization of Organic Acid Producing Aspergillus tubingensis FMS1 and its Role in Metals Leaching from Soil. Geomicrobiology Journal, 2019, 37(4): 336-344. https://doi.org/10.1080/01490451.2019, 1701585. (IF: 2.308)

 

68      Haq A, Siddiqi M, Batool S.Z, Islam A, Khan A, Khan D, Khan S, Khan H, Shah AA, Hasan F, Ahmed S and Badshah M. Comprehensive investigation on the synergistic antibacterial activities of Jatropha curcas pressed cake and seed oil in combination with antibiotics. AMB Express, 2019, 17:9(1):67. (IF: 3.298)

 

69      Khan A. Rahman U, Sidiqqui S, Irfan M, Shah AA, Badshah M, Hasan F and Khan S. Preparation and characterization of resistant starch type III from enzymatically hydrolyzed maize flour. Molecular Biology Reports, 2019, 46(4) 4565-4580. (IF: 2.316)

 

70      Rana Q, Siddiqi M, Batool S.Z, Islam A, Khan A, Khan D, Khan S, Khan H, Shah AA, Hasan F, Ahmed S and Badshah M. Bio-catalytic transesterification of mustard oil for biodiesel production. Biofuels, 2019. https://doi.org/10.1080/17597269.2019.1655907. (IF: 2.956)

 

71      Ullah S, Irfan M, Sajjad W, Rana Q, Hasan F, Khan S, Badshah M and Shah AA. Production of an alkali-stable xylanase from Bacillus pumilus K22 and its application in tomato juice clarification. Food Biotechnology, 2019, 33(4) 353-372. (IF: 1.564)

 

72      Rana Q, Rehman M.L, Irfan M, Ahmed S, Hasan F, Shah A A, Khan S and Badshah M. Lipolytic Bacterial Strains Mediated Transesterification of Non-Edible Plant Oils for Generation of High Quality Biodiesel. Journal of Bioscience and Bioengineering, 2019, 127(5): (609-617). (IF: 2.894)

 

73      Qindeel M, Ahmed N, Sabir F, Khan S and Rehman A. Development of novel pH sensitive nanoparticles loaded hydrogel for transdermal drug delivery. Drug Development and Industrial Pharmacy, 2019, 45(4):629-641. (IF: 3.225)

 

74      Sajjad W, Khan S, Ahmad M, Rafiq M, Badshah M, Sajjad W, Zada S, Khan S, Hasan F and Shah A.A. Anaerobic Digestate: A Sustainable Source of Bio-fertilizer of Ultra-Violet Radiations on cellular proteinss and Lipids of Radio Resistant Bacteria Isolated from Desert Soil. Folia Biologica (Krakow), 2018, (66): No.1. (IF: 0.432).

 

75       Irfan M,  Gonzalez CF, Raza S, Rafiq M, Hasan F, Khan S and Shah A.A. Improvement in thermostability of Xylanase from Geobacillus thermodenitrificans C5 by site directed mutagenesis. Enzyme and Microbial Technology, 2018, 111:38-47. (IF: 3.493).

 

76      Hashmi M, Khan K T, Khan W, Hasan F, Hameed A, Khan S, Badshah M and Shah A.A. Comparison Between a Newly Isolated Yeast Strain and Lalvin EC-1118 for Enhanced Ethanol Yield from Sugarcane Molasses Employing Batch and Modified Fed-Batch Fermentation. Journal of Biobased Material and Bioenergy, 2018, 12:134–142. (IF: 0.708).

 

77      Saboor A, Khan S, Shah A.A, Hasan F, Khan H and Badshah M.  Enhancement of biomethane production from cattle manure with codigestion of dilute acid pretreated lignocellulosic biomass. International Journal of Green Energy, 2017, 14(7):632-637. (IF: 2.459).

 

78      Irfan M, Tayyab A, Hasan F, Khan S, Badshah M and Shah A.A. Production and Characterization of Organic-Solvent-Tolerant Cellulase from Bacillus amyloliquefaciens AK9 Isolated from Hot Spring. Applied Biochemistry and Biotechnology, 2017, 182(4):1390-1402. (IF: 2.926).

 

79       Khan S, Kulkarni T.S, Villagomez R, Mahmood T, Lindahl S, Logan T. D, Linares-Pastén J.A and Karlsson E.N.  Crystal structure of β-glucosidase 1A from Thermotoga neapolitana and comparison of active site mutants for hydrolysis of flavonoid glucosides. Proteins: Structure, Function and Bioinformatics, 2017, 85(5):872–884. (IF: 3.756).

 

80      Shahzad H, Zainab K, Mahmood S, Malik F, Riaz H, Raza S.A and Khan S. Microbial and chemical analysis of illicit drug samples confiscated from different areas of Pakistan. Pakistan Journal of Pharmaceutical Sciences, 2016, 29(5):1639-1648. (IF: 0.684).

 

81      Khan R A, Alkreathy H.A, Shah A.S, Ahmed M, and Khan S. Protective effects of Trifolium alexandrinum L. against lung injury induced by environmental toxin CCl4 in experimental rats. Food & Nutrition Research, 2016, Article 30433. (IF: 3.894).

 

82      Sajjad W, Mehmood T.B, Hasan F, Khan S, Badshah M, Naseem A.A and Shah A.A. Characterization of Sulfur-Oxidizing Bacteria Isolated from Acid Mine Drainage and Black Shale Samples. Pakistan Journal of Botany, 2016, 48(3):1253-1262. (IF: 0.972).

 

83      Shah A.A, Nawaz A, Kanwal L, Hasan A, Khan S and Badshah M.   Degradation of poly (ε-caprolactone) by a thermophilic bacterium Ralstonia sp. strain MRL-TL isolated from hot spring. International Biodeterioration and Biodegradation-Journal, 2015, 98:35-42 (IF: 4.320).

 

84      Shah A.S, Ahmed M, Alkreathy H.A, Khan M.R, Khan R.A and Khan S. Phytochemical screening and protective effects of Trifolium alexandrinum (L.) against free radical induced stress in rats. Journal of Food Science and  Nutrition 2014, 2:6 (751–757). (IF: 2.867).

 

85      Akbar S, Hasan F, Nadhman A, Khan S and Shah A.A. Production and Purification of Poly (3-hydroxy butyrate-co-3-hydroxyvalerate) Degrading Enzyme from Streptomyces sp. AF-111. Journal of Polymers and the Environment, 2013, 21:1109–1116. (IF: 3.667).

 

86      Lindahl S, Liu J, Khan S, Karlsson E.N and Turner C. An on-line method for pressurised hot water extraction and enzymatic hydrolysis of quercetin glucosides from onions. Analytica Chimica Acta, 2013, 785 (27):50-59. (IF: 6.558).

 

87      Khan S, Lindahl, S, Turner C, and Karlsson, E.N.  Immobilization of thermostable β-glucosidase variants on acrylic supports for biocatalytic processes in hot water. Journal of Molecular Catalysis B-Enzymatic, 2012, 80:28–38. (IF: 2.269).

 

88      Khan S, Pozzo T, Megyeri M, Lindahl S, Sundin A, Turner C and Karlsson E.N. Aglycone specificity of Thermotoga neapolitana β-glucosidase 1A modified by mutagenesis, leading to increased catalytic efficiency in quercetin-3-glucoside hydrolysis. BMC Biochemistry, 2011, 12:11. (IF: 4.059).

 

89      Lindahl S, Ekman A, Khan S, Wennerberg C, Börjesson P, Sjöberg P.J.R, Karlsson E.N and Turner C. Exploring the possibility of using a thermostable mutant of β -glucosidase for rapid hydrolysis of quercetin glucosides in hot water. Green Chemistry, 2010, 12(1):159-169. (IF: 10.182).

 

90      Hasan F, Khan S, Shah A.A and Hameed A. Production of Antimicrobial Compounds by Free and Immobilized Bacillus Pumilus SAF1. Pakistan Journal of Botany, 2009, 41(3):1499-1510. (IF: 0.972).

 

91      Subhan M, Syed Besma S, Yasmeen A, Khan S, Macreadie I , Faryal R. Isolation and characterization of lovastatin producing fungi; investigating the antimicrobial and extracellular enzymatic activities. IJB 2017, 10(2), 12-20. (IF: 0.553).

 

92      Hassan M, Mehmood S, Khan N, Khan A, Khan R and Khan S. Antioxidant evaluation of methanolic extract of Tamarix aphylla and Euphorbia helioscopia. IJAR.2014, 2: 9 (62-65).

 

93      Kalsoom F, Anjum AR, Anam S, Khan S and Hasan F. Effect of temperature, pH and metal ions on amylase produced from selected indigenous extremophile bacteria in Pakistan. IJB 2018, 13(3), 262-275. (IF: 0.553).

 


 

 

1         Saleh Zada N, Rahman U, Belduz A.O, Badshah M,  Shah AA and Khan S. Designing Next-Generation  β-Glucosidases for Sustainable Biofuels, Food  and Pharma. 2nd International Conference on the Future of Biotechnology & Bioengineering (ICFBB-2025) organized by Department of Biotechnology, Kohat University of Science and Technology (KUST), June 18-19, 2025, Kohat, Pakistan. (Invited Speaker)

 

2         Khan S. Collection and characterization of metabolites of mushrooms from Azad Jammu and Kashmir, Pakistan. International Scientific and Technical Conference: Actual Problems of the Chemistry of Natural Products, organized by Academy of Sciences of the Republic of Uzbekistan, S.Yu.Yunusov Institute of the Chemistry of Plant Substances, September 19-20, 2024, Tashkent, Uzbekistan. (Plenary Presentations)

 

3         Khan S, Nawaz A, Khalid N.A, Shahzadi M, Badshah M, Shah A.A. Characterization of ES10 lytic bacteriophage isolated from hospital waste against multidrug-resistant uropathogenic E. coli. ASM Microbe 2024 meeting, June 13–17, 2024, Atlanta, Georgia, USA. (Poster Presentation)

 

4         Khan S, Sharif M, Saeeda U.H and Nawaz A. Bioactivity assessment and metabolic profiling of Phellinus gilvus. A pathway to novel therapeutics.  International Conference оn Medical Plants аnd Natural Drug Research (ICMPNDR-2024), Al Farabi Kazakh National University, June 5-7, Almaty, Kazakhstan. (Oral Presentation)

 

5         Sharif M, Fatima U, Ejaz M and Khan S. Characterization and Biological Potential of Ganoderma gibbosum collected from AJ&K, Pakistan.  International Conference оn Medical Plants аnd Natural Drug Research (ICMPNDR-2024), Al Farabi Kazakh National University, June 5-7, Almaty, Kazakhstan. (Oral Presentation).

 

6         Khan S. International Forum on Innovative Development of Biomanufacturing. 9-11 November 2023, Crown Plaza, Tianjin, China, Organized by Tianjin Institute of Industrial Biotechnology (TIB) and Chinese Academy of Sciences (CAS), Tianjin, China. (Participated Only)

 

7         Saleh Zada N, Belduz A.O, Güler H.I, and Khan S. Cloning, functional expression and characterization of active site mutants of β-glucosidse from Anoxybacillus ayderensis A9 for hydrolysis of pNPG and polydatin. International Conference and Exhibition for Science (ICES2023), February 06-08, 2023,  King Saud University, Riyadh, Saudi Arabia. (Oral Presentation)

 

8         Khan S. Biosafety Association of Central Asia and the Caucasus Conference: COVID19 - Lessons Learned 3 - 7 October 2022, Tbilisi, Georgia. Organized by Biosafety Association of Central Asia and the Caucasus in collaboration with EU CBRN Centres of Excellence Initiative, US Department of State Biosecurity Engagement Program and National Center for Disease Control and Public Health. (Participated Only)

 

9         Khan S. Metabolic Engineering Summit (MES 2019) 20-22 October 2019. Organized by International Metabolic Engineering Society (IMES) Tianjin Institute of Industrial Biotechnology, CAS. (Participated Only)

 

10       Khan A, Rahman U, Siddiqui S, Bibi A, Badshah M, Belduz A.O and Khan S. Green Synthesis of Resistant Starch type III from Maize flour and its application as drug carrier to colon. Eurasian Congress on Molecular Biotechnology at Karadeniz Technical University, 19-21 September 2019 Trabzon Turkey. (Oral Presentation)

 

11      Marium S, Farooq S, Belduz A.O, Badshah M, Shah AA and Khan S. Exploitation of keratinase potential for the green Synthesis of silver nanoparticles. Molecular Biotechnology at Karadeniz Technical University, S 19-21 eptember, 2019 Trabzon Turkey. (Oral Presentation)

 

12      Salehzada N, Hameed F, Bari F and Khan S. Phenotypic detection of Mettalo-β-Lactamase (MBL) in Imipenem Resistant Pseudomonas aeruginosa, a study from a tertiary care hospital in Peshawar.Molecular Biotechnology at Karadeniz Technical University, 19-21 September 2019 Trabzon Turkey. (Oral Presentation)

 

13      Khan S.I, Zarin A, Belduz A.O, Canakci S, Ahmed S, Hasan F, Khan S and Shah AA. Phenotypic detection of Mettalo-β-Lactamase (MBL) in Imipenem Resistant Pseudomonas aeruginosa, a study from a tertiary care hospital in Peshawar. Molecular Biotechnology at Karadeniz Technical University, 19-21, September 2019 Trabzon Turkey. (Oral Presentation)

 

14      Siddiqui S, Khan A, Rahman U and Khan S. In vitro digestibility and Physicochemical properties of Enzymatically Produced Resistant Starch from Maize. 12th International Biennial Conference Pakistan Society of Microbiology on Next Generation of Microbial Therapeutics at Abbottabad University of Science and Technology 20-22 February 2019 Abbottabad, Pakistan.  (Oral Presentation)

 

15      Bibi A, Khan A, Rahman U and Khan S. Synthesis and Characterization of Enzymatically Produced Resistant Starch Coated Drug Microspheres and their Antimicrobial Potential.  12th International Biennial Conference Pakistan Society of Microbiology on Next Generation of Microbial Therapeutics at Abbottabad University of Science and Technology 20-22 February 2019 Abbottabad, Pakistan.  (Oral Presentation)

 

16      Ali H, Bibi A, Khan A, Badshah M, Shah AA and Khan S. Prebiotic potential of enzymatically prepared resistant starch type III and its effect on controlling metabolic dysfunction on rate physiology.  International Pharmacy Conference and Exhibition (IPCE) on “Emerging Trends in Drug Development, Therapeutics and Commercialization” at Department of Pharmacy, University of Swabi, 18-20 June 2019, Swabi, KPK, Pakistan. (Oral Presentation)

 

17      Alam T, Sajjad W, Hasan F, Khan S and Ali Shah A. Evaluation of Antioxidant Potentials of Intracellular Bioactive Metabolites of UV Resistant Streptomyces chilikenes TMM1 Isolated from Thal Desert, Pakistan. 12th International Biennial Conference Pakistan Society of Microbiology on Next Generation of Microbial Therapeutics at Abbottabad University of Science and Technology 20-22 February 2019 Abbottabad, Pakistan. (Poster Presented)

 

18      Salahuddin A, Javeria K, Irfan M, Hasan F, Khan S and Ali Shah A. Immobilization of Partially Purified β-1-4-Xylanase Isolated from Thermomyces lanuginosus. 12th International Biennial Conference Pakistan Society of Microbiology on Next Generation of Microbial Therapeutics at Abbottabad University of Science and Technology 20-22 February 2019 Abbottabad, Pakistan. (Poster Presented)

 

19      Khan S. Development of microbial enzymes for modification of polyphenolic glucosides. 2018 Synthetic Biotechnology Training Course for   Developing Countries, jointly 1st TWAS Symposium on Plant Natural Product Synthesis by Microbes held on 15-19 January 2018, in Beijing, China. (Oral Presentation)

 

20      Khan A, Bano K, F Kalsoom, Hasan F, Shah A A and Khan S. Rheinheimera TS-5 isolated as a potential producer of polyextremophilic α-amylase. 10th Biennial Conference of Pakistan Society of Microbiology, 25-28 March 2016, at Department of Microbiology and Molecular Genetics, University of the Punjab Lahore. (Poster Presented)

 

21      Haq A, Badshah M and Khan S. Comparison of antimicrobial activities of different parts of geographically two different species of Jatropha curcus Linn fruit. 1st international conference on recent innovations in pharmaceutical sciences (ICRIPS-2015), 3-5 March, 2015, Pakistan. (Poster presented).

 

22      Turner C, Börjesson P, M.Co, Ekman A, Khan S, Samuelsson J, Lindahl S, Mijangos Trevino L, Mustafa A and Karlsson E.N. Water, ethanol and supercritical carbon dioxide – are these solvents really green? Gordon Green Chemistry Conference, 22-27 July 2012, Lucca, Italy. (Poster Presented)

 

23      Lindahl S, Liu J, Khan S, Karlsson E.N and Turner C. On-line extraction and hydrolysis of polyphnolic glucosides. 2nd International Symposium on Hyphenated Techniques for Sample Preparation. January 31-February 1, 2012, Bruges, Belgium. (Poster Presented)

 

24      Khan S, Mahmood Tahir, Lindahl S, Turner C and Nordberg Karlsson E. Mutagenesis of substrate interacting residues in the active site of TnBgl1A for improved deglycosylation of antioxidants. 16th European Carbohydrate Symposium 3-7 July 2011, Naples, Italy. (Poster Presented)

 

25      Khan S, Lindahl S, Turner C and Karlsson E.N. Immobilization of thermostable β-glucosidases (TnBgl1A) for modification of antioxidants at high temperature. 9th Carbohydrate Bioengineering Meeting 15-18 May 2011, Lisbon, Portugal. (Poster Presented)

 

26      Karlsson E.N, Khan S, Pozzo T, Mahmood T and Gulshan K.Z. Mutagenesis of substrate interacting residues in the active site of TnBgl1A for modification of flavonoid antioxidants. 9th Carbohydrate Bioengineering Meeting, 15-18 May 2011, Lisbon, Portugal. (Poster Presented)

 

27      Khan S, Lindahl S, Turner C and Karlsson E.N. Immobilization of thermostable β-glucosidases TnBglA for modification of antioxidants using hot water. Thermophiles 16-21 August 2009, Beijing, China. (Poster Presented)


1.       Principal Investigator in a project (PKR. 12.856022 million) funded by Higher Education Commission of Pakistan “Bacteriophage Derived Proteins a Novel Approach as Enzobiotics to Combat Multi Drug Resistance Bacteria”

 

2.       Principal Investigator in a project (PKR. 8.954,722 million) funded by Pakistan Science Foundation and TUBITAK, Turkey “Specific designing of β-glucosidase for modification of natural products with the aim to enhance their cancer preventing potential”

 

3.       Principal Investigator in a project (PKR. 3.807,224 million) funded by Higher Education Commission of Pakistan “Utilization of poultry feather waste by keratinases producing microorganisms for production of sustainable value-added products and biogas”

 

4.       Principal Investigator in a project (PKR. 2.68 million) funded by Higher Education Commission of Pakistan and Turkey “Green and Economical Production of Resistant Starch”

 

5.       Principal Investigator in a project (PKR. 0.5 million) funded by Higher Education Commission of Pakistan “Isolation and molecular identification of amylase producing bacterial strains and characterization of amylase”

 

6.       Co-principal Investigator in a project (PKR 9.876 million) funded Higher Education Commission of Pakistan “Synthesis of biowaste derived titania Nanoparticles and their efficiency in developing Novel restorative material for dental caries treatment

 

7.       Co-principal Investigator in a project (PKR 4.08 million) funded Higher Education Commission of Pakistan “Utilization of Jatropha curcas for coproduction of biodiesel, biogas, biofertilizer and antimicrobials”

 

8.       Co-principal Investigator in a project in a project (PKR 3.8 million) funded by International Foundation for Science “Bacteriophages Proteins: Alternative Approach to Antibiotics against Gram Negative Bacteria in Resistance Era”

 



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