Name: Dr. Awais Rasheed
Designation: Assistant Professor
Department: Plant Sciences


  • PhD (QAU)
  • Postdoc (CIMMYT-China)
  • Expert of State Administration of Foreign Experts Affairs P.R. China (A Level).
  • Vavilov-Frankel Fellow (VFF-2013)  by Bioversity International (Italy) and GRDC (Australia). 

Phone: +92-51 9064-3137
Status: On Job
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Research Interests Publications Conferences Research Projects

High-throughput phenotyping
Molecular Breeding

1)    Hao Y, Rasheed A, Zhu Z, Wulff B, He ZH (2019) Harnessing wheat Fhb1 for Fusarium resistance. Trends in Plant Science doi: 10.1016/j.tplants.2019.10.006

2)    Li J, Chen GB, Rasheed A, Li D, Sonder K, Espinosa CZ, Wang J, Costich DE, Schnable PS, Hearne SJ, Li H (2019). Identifying loci with breeding potential across temperate and tropical adaptation via EigenGWAS and EnvGWAS. Molecular Ecology 28(15): 3544-3560 doi: 10.111/mec.15169

3)    He ZH, Xia XC, Zhang Y, Zhang Y, Xiao Y, Chen X, Li S, Hao Y, Rasheed A, Xin Z, Zhuang Q, Yang E, Fan Z, Yan J, Sing RP, Braun J-C (2019). China-CIMMYT collaboration enhances wheat improvement in China. Frontiers of Agricultural Science and Engineering 6(3): 233-239 doi: 10.15302/J-FASE-2019254

4)    Khalid M, Afzal F, Gul A, Amir R, Ahmed Z, Subhani A, Xia XC, He ZH, Rasheed A (2019) Molecular characterization of 87 functional genes in wheat diversity panel and their association with phenotypes under well-watered and water-limited conditions. Frontiers in Plant Science 10: 717 doi: 10.3389/fpls.2019.00717

5)    Liu J, Rasheed A, He ZH, Imtiaz M, Arif A, Mahmood T, Ghafoor A, Siddiqui S, Ilyas MK, Wen W, Gao F, Xie C, Xia XC (2019) Genome-wide variation patterns between landraces and cultivars uncover divergent selection during modern wheat breeding. Theoretical and Applied Genetics 132 (9): 2509-2523 doi: 10.1007/s00122-019-03367-4

6)    Li F, Wen W, Liu JD, Zhang Y, Cao S, He ZH, Rasheed A, Jin H, Zhang C, Yan J, Zhang P, Wan Y, Xia XC (2019) Genetic architecture of grain yield in bread wheat based on genome-wide association studies. BMC Plant Biology 19: 168 doi: 10.1186/s12870-019-1781-3

7)    Afzal F, Li H, Gul A, Subhani A, Ali A, Mujeeb-Kazi A, Ogbonnaya F, Trethowan R, Xia XC, He ZH, Rasheed A (2019) Genome-Wide Analyses Reveal Footprints of Divergent Selection and Drought Adaptive Traits in Synthetic-Derived Wheats. G3: Genes, Genomes, Genetics 9(6): 1957-1973 doi: 10.1534/g3.119.400010

8)    Hassan A, Mengjiao Y, Fu L, Rasheed A, Zheng B, Xia XC, Xiao XY, He ZH (2019) Accuracy assessment of plant height using unmanned aerial vehicle for quantitative genomic analysis in bread wheat. Plant Methods 15: 37 doi: 10.1186/s13007-019-0419-7

9)    Mujeeb-Kazi A, Munns R, Rasheed A, Ogbonnaya F, Ali N, Hollington P, Dundas I, Saeed N, Wang R, Rengasamy P, Saddiq MS, Leon JL, Ashraf M, Rajaram S (2019) Breeding strategies for structuring salinity tolerance in wheat. Advances in Agronomy 155: 121-187 doi: 10.1016/bs.agron.2019.01.005

10) Rasheed A, Xia XC (2018) From molecular markers to genome-based breeding in wheat. Theoretical and Applied Genetics 132(3): 767-784 doi: 10.1007/s00122-019-03286-4

11) Rasheed A, Jin H, Xiao YG, Zhang Y, Hao Y, Zhang Y, Hickey L, Morgounov A, Xia XC, He ZH (2019) Allelic effects and variations for key bread-making quality genes in bread wheat using high-throughput molecular markers. Journal of Cereal Science 85(1): 305-309 doi: 10.1016/j.jcs.2018.12.004

12) Hassan MA, Yang M, Rasheed A, Yang G, Reynolds M, Xia XC, Xiao YG, He ZH (2018) A rapid monitoring of NDVI across the wheat growth cycle for grain yield prediction using a multi-spectral UAV platform. Plant Science 282(5): 95-103doi: 10.1016/j.plantsci.2018.10.022

13) Hassan MA, Yang M, Rasheed A, Jin X, Xia XC, Xiao YG, He ZH (2018) Time-series multispectral indices from unmanned aerial imagery reveal senescence rate in bread wheat. Remote Sensing 10: 809 doi: 10.3390/rs10060809

14) Khalid M, Gul A, Amir R, Ali M, Afzal F, Quraishi UM, Ahmed Z, Rasheed A (2018) QTL mapping for seedling morphology under drought stress in wheat cross synthetic (W7984)/Opata. Plant Genetic Resources 16(4): 359-366 doi: 10.1017/S1479262118000023

15) Rasheed A, Ogbonnaya F, Lagudah E, Appels R, He ZH (2018) Goatgrass genome diversity for wheat improvement. Nature Plants 4(2): 56-58 doi: 10.1038/s41477-018-0105-1

16) Huihui L, Rasheed A*, Hickey L, ZH He (2018) Fast-forwarding genetic gain. Trends in Plant Sciences 23(3): 184-186 doi: 10.1016/j.tplants.2018.01.007

17) Liu JD, He ZH, Rasheed A, Weie W, Yan J, Zhang P, Wan Y, Zhang Y, Xie C, Xia XC (2017) Genome-wide association mapping of black point reaction in common wheat (Triticum aestivum L.). BMC Plant Biology 17: 220. doi: 10.1186/s12870-017-1167-3

18) Almas F, Hassan MA, Bibi A, Ali M, Lateef S, Mahmood T, Rasheed A, Quraishi UM (2017) Identification of genome-wide single-nucleotide polymorphisms associated with tolerance to chromium toxicity in spring wheat (Triticum aestivum L.). Plant and Soil 422(1-2) 371-384 doi: 10.1007/s11104-017-3436-1

19) Rasheed A, Mujeeb-Kazi A, Ogbonnaya FC, He ZH, Rajaram S (2017) Wheat genetic resources in post-genomics era: promise and challenges. Annals of Botany 121(4): 603-616 doi: 10.1093/aob/mcx148

20) Aziz A, Mahmood T, Shazadi K, Mahmood Z, Mujeeb-Kazi A, Rasheed A* (2017) Genotypic variation and genotype x environment interaction for yield-related traits in synthetic hexaploid wheats under a range of optimal and heat-stressed environments. Crop Science 58(1): 295-303 doi: 10.2135/cropsci2017.01.0035

21) Li J, Rasheed A, Guo Q, Dong Y, Liu JD, Xia XC, Zhang Y, He ZH (2017) Genome-wide association mapping of starch granule size distribution in common wheat. Journal of Cereal Science 77(9): 211-218 doi: 10.1016/j.jcs.2017.08.016

22) Rasheed A, Hao YF, Xu YB, Xia XC, Varshney RK, He ZH (2017) Crop breeding chips and genotyping platforms: progress, challenges and perspectives. Molecular Plant 10(8): 1047-1064. doi: 10.1016/j.molp.2017.06.008

23) Ogbonnaya F, Rasheed A, Emeka O, Jighly A, Taddesee W (2017) Genome-wide association mapping for agronomic and physiological traits in wheat in a range of heat stress environments. Theoretical and Applied Genetics 130(9): 1819-1835. doi: 10.1007/s00122-017-2927-z

24) Afzal A, Reddy B, Gul A, Khalid M, Subhani A, Shazadi K, Quraishi UM, Ibrahim AMH, Rasheed A* (2017) Physiological, biochemical and agronomic traits associated with drought tolerance in synthetic-derived wheat diversity panel. Crop and Pasture Science 68(3): 213-224. doi: 10.1071/CP16367

25) Gao FM, Ma DY, Yin GH, Rasheed A, Dong Y, Xiao YG, Xia XC, Wu XX, He ZH (2017) Genetic progress in grain yield and physiological traits in Chinese wheat cultivars of Southern Yellow and Huai Valley Winter Wheat Zone since 1950. Crop Science 57: 760-773. doi: 10.2135/cropsci2016.05.0362

26) Dong Y, Zhang Y, Liu JD, Rasheed A, Xiao YG, Fu LP, Yan J, Wen W, Zhang Y, Jing RL, Xia XC, He ZH (2016) Genome-wide association of stem water soluble carbohydrates in bread wheat. Plos One 11(11): e0164293 doi: 10.1371/journal.pone.0164293

27) Rasheed A, Wen W, Gao FM, Zhai SN, Jin H, Liu JD, Guo Q, Zhang YJ, Dreisigacker S, Xia XC, He ZH (2016) Development and validation of high-throughput KASP assays for functional genes in wheat. Theoretical and Applied Genetics 129(10): 1843-1860 doi: 10.1007/s00122-016-2743-x

28) Ilyas M, Rasheed A, Mahmood T (2016) Functional characterization of germin and germin-like protein genes in various plant species using transgenic approaches. Biotechnology Letters 38(9):1405-1421 doi: 10.1007/s10529-016-2129-9

29)   Hanif M, Gao FM, Liu JD, Wen WE, Zhang YJ, Rasheed A, Xia XC, He ZH, Cao SH (2016) TaTGW6-A1, an ortholog of rice TGW6, is associated with grain weight and yield in bread wheat. Molecular Breeding 36:1 doi: 10.1007/s11032-015-0425-z

30) Rasheed A, Mahmood T, Xia XC, Bux H, Quraishi UM, Mahmood Z, Mirza JI, Mujeeb-Kazi A, He ZH (2016) Comparison of alleles at economically important loci in wheat landraces and cultivars from Pakistan. Crop Science 56(1): 287-301 doi:10.2135/cropsci2015.01.0015

31) Khan Z, Qazi J, Rasheed A, Mujeeb-Kazi A (2016) Diversity in D genome synthetic hexaploid wheat association panel for seedling emergence traits under salinity stress. Plant Genetic Resources doi: 10.1017/S1479262116000198

32) Gao FM, Liu JD, Wen WE, Rasheed A, Xia XC, He ZH (2015) QTL mapping for agronomic traits in Zhonghou8592B/Chinese Spring population. Frontiers in Plant Science 6:1099. doi: 10.3389/fpls.2015.01099

33) Ain Q, Rasheed A, Anwar A, Mahmood T, Mahmood T, Xia XC, He ZH, Mujeeb-Kazi A, Quraishi UM (2015) Genome-wide association analysis for grain yield in historical wheat cultivars from Pakistan under rainfed conditions. Frontiers in Plant Science 6:743 doi: 10.3389/fpls.2015.00743

34) Ali I, Sardar Z, Rasheed A, Mahmood T (2015) Molecular characterization of the puroindoline-a and b alleles in synthetic hexaploid wheats and in silico functional and structural insights into Pina-D1. Journal of Theoretical Biology 376(7): 107. doi: 10.1016/j.jtbi.2015.04.001

35) Ali A, Arshad M, Naqvi SMS, Sher H, Kazi AG, Mujeeb-Kazi A, Rasheed A* (2015) Comparative assessment of synthetic-derived and conventional bread wheat advanced lines under osmotic stress and implications of molecular analysis. Plant Molecular Biology Reporter 33(6): 1907-1917 doi:10.1007/s11105-015-0884-8

36) Ilyas M, Mahmood T, Ali A, Babar M, Mujeeb-Kazi A, Rasheed A* (2015) Characterization of D genome diversity for tolerance to boron toxicity in synthetic hexaploid wheat and in silico analysis of candidate genes. Acta Physiologiae Plantarum 37(2): 17. doi: 10.1007/s11738-014-1765-8

37) Zegeye H, Rasheed A*, Makdis F, Badebo A, Ogbonnaya FC (2014) Genome-wide association for stripe rust resistance at seedling and adult plant stages in synthetic hexaploids. PLoS One 9(8): e105593 doi: 10.1371/journal.pone.0105593

38) Ali A, Arshad M, Naqvi SM, Fatima S, Sher H, Kazi AG, Rasheed A*, Mujeeb-Kazi A (2014) Exploitation of synthetic derived wheats through osmotic stress responses for drought tolerance improvement. Acta Physiologiae Plantarum 36(9): 2453-2465 doi: 10.1007/s11738-014-1618-5

39) Hanif U, Rasheed A, Kazi AG, Munir M, Mujeeb-Kazi A (2014) Analysis of genetic diversity in synthetic wheat assemblage (triticum turgidum × Aegilops tauschii; 2n=6x=42; aABBDD) for winter wheat breeding. Cytologia 79(4): 485-500 doi: 10.1508/cytologia.79.485

40) Rasheed A, Xia XC, Ogbonnaya FC, Mahmood T, Zhang Z, Mujeeb-Kazi A, He ZH (2014) Genome-wide association for grain morphology in synthetic hexaploid wheats using digital imaging analysis. BMC Plant Biology 14: 128 doi: 10.1186/1471-2229-14-128

41) Rasheed A, Xia XC, Mahmood T, Yan YM, Appels R, He ZH (2014) Wheat storage proteins: recent advances in molecular genetics, diversity and breeding applications. Journal of Cereal Science 60(1): 11-24 doi: 10.1016/j.jcs.2014.01.020

42) Sania A, Rasheed A*, Bux H, Kazi AG, Rauf A, Mahmood T, Mujeeb-Kazi A (2014) Stripe rust resistance in Triticum durum - T. monococcum and T. durum - T. urartu amphiploids. Australasian Plant Pathology 43: 109-113 doi: 10.1007/s13313-013-0237-8

43) Mujeeb-Kazi A, Kazi AG, Dundas I, Rasheed A, Ogbonnaya FC, Bonnett D, Kishii M, Wang RC, Xu SS, Peidu C, Mahmood T, Bux H, Rizwan S. (2013) Genetic diversity for wheat improvement as a conduit to food security. Advances in Agronomy 122: 179-257. doi: 10.1016/B978-0-12-417187-9.00004-8

44) Rasheed A*, Safdar T, Kazi AG, Akram Z, Mahmood T, Mujeeb-Kazi A (2012) Characterization of HMW-GS and evaluation of their diversity in Elite synthetic hexaploid wheats. Breeding Science 62: 365-370 doi: 10.1270/jsbbs.62.365

45) Rafique K, Rasheed A*, Kazi AG, Bux H, Naz F, Mahmood T, Mujeeb-Kazi A (2012) Powdery mildew resistance in some new wheat amphiploids (2n = 6x = 42) derived from A- and S-genome diploid progenitors. Plant Genetic Resources 10(3): 165-170. doi: 10.1017/S1479262112000202

46) Bibi A, Ajmal S, Rasheed A*, Kazi AG, Mujeeb-Kazi A (2012) High molecular weight (HMW) glutenin subunit compositions of the Elite-II synthetic hexaploid wheat sub-set (Triticum turgidum × Aegilops tauschii; 2n=6x=42; AABBDD). Plant Genetic Resources. 10(1): 1-4. doi: 10.1017/S1479262111000888


Books/Book Chapters

1)      Borner A, Ogbonnaya FC, Roder M, Rasheed A, Periyannan S, Lagudah E (2015) Aegilops tauschii introgressions in wheat. In: Molnar-Lang M, Ceoloni C, Dolezel J (Eds.). Alien introgressions in wheat. Pp 245-271. Springer, New York, USA. ISBN: 978-3-319-23494-6. doi: 10.1007/978-3-319-23494-6_10

2)    Kazi AG, Rasheed A, Mujeeb-Kazi A. 2012. Biotic stress and crop Improvement: A wheat focus around novel strategies. In: Environmental challenges and crop improvement (Ozturk M. ed.). Springer New York. ISBN:

3)    Rashhed A*, Gul-Kazi A, Mujeeb-Kazi A. 2012. An overview of Omics to improve wheat grain quality. In: Crop Improvement: New approaches and modern techniques. Modern techniques in crop improvement (Hakeem KR, Ozturk M, Ahmed P. eds.). Springer New York. ISBN 978-1-4614-7027-4. doi: 10.1007/978-1-4614-7028-1_10         

4)    Bux H, Ashraf M, Rasheed A, Dipak SP, Kazi AG, Afzaal M. 2012. Molecular basis of disease resistance in cereal crops: An overeview. In: Crop production for agriculture improvement. Ashraf, M.; öztürk, M.; Ahmad, M.S.A.; Aksoy, A. (Eds.). Springer. ISBN 978-94-007-4115-7. doi: 10.1007/978-94-007-4116-4_18. pp 477-489

·      ·      2019-2021: Molecular mapping and map-based cloning for disease resistance genes and development of disease resistant wheat germplasm. PSF-NSFC Joint Collaborative Project between Pakistan and China. Amount: 14 million PKR + 2 million RMB

·      2019: Optimizing Unmanned Aerial Vehicle (UAV) based high-throughput phenotyping for physiology, grain yield and disease prediction in wheat. USAID/AIP/CIMMYT. Amount: 9000 US$

·      Capturing novel diversity for earliness per se and grain weight in D-genome synthetic hexaploid wheats under climate change scenario. National Natural Science Foundation (NNSF) China. Amount: ~70,000 US$. (Principal Investigator). Completed

·      Development of nucleotide and digital barcodes for historical wheat cultivars for varietal identification and digital repository. US-AID’s Agriculture Innovation Program (AIP), Pakistan. Amount: 30,000 US$. (Principal Investigator). Completed

Molecular and phenomic characterization of synthetic hexaploids to enhance its breeding value for grain size and weight. Bioversity International, Italy. Amount: 20,000 US$. (Principal Investigator). Completed.

Quaid-i-Azam University Islamabad, 45320, Pakistan.
Tel: +92-51 9064 0000,